A Kinematic Comparison of Running, Standing Cycling, and Seated Cycling
Maggie Davis and Erin Abercrombie
Biomechanics Project
November 20, 2001

TABLE OF CONTENTS

Introduction/Lit Review

Hypotheses

Methods

Results

Discussion

References

 

 

INTRODUCTION

Popular opinion suggests that triathletes should stand at the end of the cycle portion of a race in an effort to ease the cycle to run transition. Authorities advise, "During the last couple of miles of the bike portion, it is suggested to ride out of the saddle to increase the blood flow to your running muscles. You are in effect simulating the running motion and preparing your legs for a smooth and fast transition into the running leg of the event" (Hobson, Campbell, & Vickers, 2001). No empirical evidence of kinematic data exists in the literature to support this recommendation to the best of our knowledge. Therefore, the purpose of our study is to compare the kinematics of seated and standing cycling to the kinematics of running. We will examine the hip, knee, and ankle joints of the lower body, and we will quantify their changes in position and velocity with respect to time. We will also determine cycle and running frequencies.
In order to draw our hypotheses, we will review relevant literature pertaining to the sports of running and cycling. To begin, running gait begins when one foot contacts the ground and ends when the same foot contacts the ground again (Novacheck, 1998). The stance/support phase, starting with heel strike and ending with toe off, occurs when one foot is in contact with the ground (Adelaar, 1986; Novacheck, 1998). The float/unsupport phase occurs when the body is in the air (Adelaar, 1986; Novacheck, 1998). The swing phase, starting with toe off and ending with heel strike, occurs when the leg is moving through the air (Adelaar, 1986; Novacheck, 1998).
Maximum flexion and extension angles of the lower extremity joints during running are frequently reported in the literature. For example, hip joint angles in the literature include 125 degrees maximum flexion during swing phase and 185 degrees maximum extension following toe off (Novacheck, 1998), and maximum flexion of 136 degrees and maximum extension of 196 degrees (Quigley & Richards, 1996).
Knee joint angles are reported as 90 degrees maximum flexion during swing phase and 155 degrees maximum extension during toe off (Novacheck, 1998), and maximum knee flexion of 92.6 degrees and maximum extension of 171.8 degrees (Quigley & Richards, 1996). Other sources provide average maximum knee flexions of 94.2 degrees (Hausswirth, Bigard, & Guezennec, 1997) and 120 degrees (Herzog, Guimaraes, Anton, & Carter-Erdman, 1991) during the non-support phase of running. Another source provides an average peak knee flexion of 133.5 degrees (Dixon, Collop, & Batt, 2000).
Ankle joint angles are reported as 60 degrees dorsiflexion during stance phase and 110 degrees maximum plantarflexion just following toe off (Novacheck, 1998), and maximum dorsiflexion of 67 degrees and maximum plantarflexion of 118.7 degrees (Quigley & Richards, 1996). Another source reports an average maximum ankle dorsiflexion of 61.1 degrees (Dixon, Collop, & Batt, 2000).
In regards to cycling, one complete circular motion is called a pedal cycle, and it consists of a power phase, beginning at top center and ending at bottom center, and a recovery phase, moving from bottom center to top center (Sanner & O'Halloran, 2000). Maximum flexion and extension joint angles of the lower extremity during sitting cycling are commonly described in the literature.
For instance, one source reports that in the early power phase the thigh begins 10-20 degrees below horizontal, the knee is flexed to 110 degrees, and the ankle is dorsiflexed at 90 degrees (Sanner & O'Halloran, 2000). In the early recovery phase the thigh is flexed to 50-75 degrees below the horizontal, the knee is flexed to 50 degrees, and the ankle is moderately plantarflexed (Sanner & O'Halloran, 2000). Additional joint angles for sitting cycling found in literature include hip angles of 57 degrees maximum flexion and 97 degrees maximum extension, knee angles of 77 degrees maximum flexion and 140 degrees maximum extension, and ankle angles of 100 degrees maximum flexion and 117 degrees maximum extension (Caldwell, Hagberg, McCole, & Li Li, 1999).
Maximum flexion and extension angles of the lower extremity joints during standing cycling are also reported in the literature, such as hip angles of 86 degrees maximum flexion and 132 degrees maximum extension, knee angles of 77 degrees maximum flexion and 157 degrees maximum extension, and ankle angles of 100 degrees maximum flexion and 128 degrees maximum extension (Caldwell, Hagberg, McCole, & Li Li, 1999).
Angular velocities of the lower extremity during running reported in the literature are maximum hip flexion velocity of 175.6 degrees/second and maximum hip extension velocity of -467.3 degrees/second (Quigley & Richards, 1996). Maximum knee positive angular velocity is 727.71 degrees/second and maximum knee negative angular velocity is -601.65 degrees/second (Creagh, Reilly, & Lees, 1998). Additional maximum knee flexion velocity values are 617.12 degrees/second (Dixon, Collop, & Batt, 2000) and 703.1 degrees/second (Quigley & Richards, 1996), while maximum knee extension velocity is reported as -421.3 degrees/second (Quigley & Richards, 1996).
Maximum ankle dorsiflexion is -464.9 degrees/second and maximum plantarflexion is reported as 873.7 degrees/second (Quigley & Richards, 1996). Another source reports dorsiflexion and plantarflexion velocity values of 464.13 degrees/second (Dixon, Collop, & Batt, 2000).
To the best of our knowledge no specific values for angular velocities of the lower extremity during seated or standing cycling exist in the literature.
Frequencies of all three conditions are commonly reported in the literature. For example, typical running frequencies include 1.0-1.5 Hz (Millet & Vleck, 1999), 2.78 + 0.05 steps/second or 1.39 + 0.02 Hz (Gottschall & Palmer, 2000), and 181.18 steps/minute or 1.51 Hz (Creagh, Reilly, & Lees, 1998). Typical seated cycling frequencies are reported as 1.5-2.0 Hz (Millet & Vleck, 1999), 82 rpm or 2.14 Hz (Caldwell, Hagberg, McCole, & Li Li, 1999), and 85.2+ 9.2 rpm or 2.23 + 0.24 Hz (Marsh & Martin, 1995). Another article states that a basic rule of cycling is being able to maintain a cadence of at least 70 rpm and preferably more than 80 rpm for a prolonged period (Sanner & O'Halloran, 2000). Typical standing cycling frequency is reported as 64 rpm or 1.68 Hz (Caldwell, Hagberg, McCole, & Li Li, 1999).
The studies that report hip, knee, and ankle angles include subjects of different sexes, ages, and training backgrounds. For example, one study used male biathletes and triathletes (Quigley & Richards, 1996), another used female middle distance runners (Dixon, Collop, & Batt, 2000), and still another used elite male cyclists (Caldwell, Hagberg, McCole, & Li Li, 1999). In addition, the studies that reported hip, knee, and ankle angles also included a range of subjects with different demographic characteristics. While one used male biathletes and triathletes (Quigley & Richards, 1996), another used female runners in their study (Creagh, Reilly, & Lees, 1998). Finally, the studies that reported running, standing cycling, and seated cycling frequencies also used different types of participants. For example, while one study used male cyclists (Marsh & Martin, 1995), another used male and female elite and junior triathletes (Millet & Vleck, 1999).
The studies that we reviewed to obtain lower extremity angles, velocities, and frequencies utilized participants with varying characteristics. Therefore, comparing the running, seated cycling, and standing cycling angles, velocities, and frequencies cited in this literature review in an effort to determine whether standing cycling is more similar to running than is seated cycling will yield inappropriate conclusions. Since no one study has compared the kinematics of running, standing cycling, and seated cycling to the best of our knowledge, our study aims to analyze joint angles, velocities, and frequencies during the three different activities to determine if scientific evidence supports popular opinion.
The literature often reports that changes in seat height or seat tube angles lead to alterations in body position, which lead to changes in lower extremity kinematics. One study reports that differences in cycling performances can be attributed to joint angle changes resulting from alterations in seat height (Too & Landwer, 2000). Adjusting the bike setting to have a higher seat height and a more forward seat tube angle causes a change in the body position from seated cycling to emulate standing cycling to a greater degree.
Many studies provide evidence that the posture adopted with a higher seat height and a more forward seat tube angle also resembles posture adopted during running. For example, one study specifically states that cyclists prefer a higher seat position because it more resembles a walking or running posture (Sanner & O'Halloran, 2000). An article concerning seat tube angles states that the further forward the seat tube angle is set, the more similar the body posture is to running, and they go on to claim that adopting this seat tube angle will minimize muscle recruitment alterations and ease the transition of cycling to running (Garside & Doran, 2000).

HYPOTHESES

The findings in the literature state that the body position of a cyclist using a higher seat height and a more forward seat tube angle, which resembles standing cycling, emulates the body position of a runner. Therefore, we hypothesize that the kinematics of standing cycling, including hip, knee, and ankle angles and velocities, along with frequencies, should be more similar to running than the kinematics of seated cycling.

METHODS

In order to gather kinematic data, a participant with experience in running and cycling was recruited. The participant was a male, age 42, with a height of six feet and a weight of 153 pounds. His race history within the last year include one triathlon (40 mile bike, 12 mile canoe, 13 mile run) and three 10K, one 12K, and four 5K running races. His weekly training schedule includes running 35 miles, biking 60 miles, and swimming 3,000 yards.
During the filming, the subject performed at an RPE of 16 to simulate the intensity of race pace. To gather cycling data a spinning bike was set up so that the subject's left side was filmed in the sagittal plane. The video camera was aligned perpendicularly to this field of view at a distance of 15 feet away from the bike. The camera was 66 centimeters above the ground to capture the lower extremities of the subject and the entire range of motion. It was leveled side-to-side and front-to-back, locked into place, and the shutter speed was set at 1/60th of a second.
A reference scale of 0.9 meters was placed in the field of view. Additional lighting was placed behind the camera and pointed towards the action as the filming took place indoors. Joint markers of different sizes and contrasting colors were placed at the trunk, the greater trochanter, the lateral condyle, the lateral malleolus, and the fifth metatarsal in order to obtain hip, knee, and ankle angles (Figure 1). The subject was also outfitted with a heart rate monitor.
After the subject positioned himself on the bike, the camera was auto focused at the place of action and manually focused on the subject ensuring that all body markers were visible. First the subject was taped while cycling sitting down for two minutes at an RPE of 16 and heart rate readings were taken at minutes one and two. The subject then transitioned into standing cycling for another two minutes at an RPE of 16. Heart rate readings were again taken at minutes one and two.
The next portion of the trials involved filming the subject running on the treadmill. The camera was set up to film the subject's left side in the sagittal plane. The video camera was then aligned perpendicularly to this field of view at a distance of 12.5 feet away from the treadmill. The camera was 89 centimeters above the ground to capture the lower extremities of the subject and the entire range of motion. It was leveled side-to-side and front-to-back, locked into place, and the shutter speed was set at 1/60th of a second.
A reference scale of 0.9 meters was placed in the field of view. Additional lighting was placed behind the camera and pointed towards the action as the filming took place indoors. Joint markers and the heart rate monitor remained intact during the filming. Again the subject was filmed running for 2 minutes at an RPE of 16, and heart rate readings were taken at one and two minutes.
The video was captured using the "Shortcut to Video Capture" program. Approximately 5 seconds of each condition was captured and made into an ".avi" file. Video editing was done using the Premiere 4.0 program. Each ".avi" file was trimmed to allow for one revolution on the bike and one stride in running with three frames before and after the desired action. The clips were finalized into movie ".avi" files.
The movies were digitized at thirty frames per second using the Video Expert program. Specific points that were used in every frame were the trunk, hip, knee, ankle, and fifth metatarsal in the sagittal plane. Finally the "x" and "y" coordinates that were obtained from digitizing were imported to Microsoft Excel where joint angles, velocities, and accelerations were obtained.

RESULTS

One complete period of seated cycling, standing cycling, and running was analyzed to determine joint angle positions, joint angular velocities, and frequencies of the movements. Joint angles and velocities for the knee, hip, and ankle were graphed as a function of the percent of the period of movement.

Heart Rate and Frequency
Table 1 lists the average heart rate for each condition, and the calculated frequency of each condition.

Joint Angle Measurements
Knee
In all three conditions, the knee joint angle exhibits the same basic trend of beginning at greatest knee extension, moving to greatest knee flexion, and ending in greatest knee extension again. The seated cycle condition had a smoother transition through the angles than the running and standing cycle conditions. The seated knee angle moved from 146.5 deg to 69.2 deg and back to 148. 3 deg. The range of motion of the knee during seated cycling was 78.1 deg. The other two conditions of standing cycle and running did not have as smooth an arc of joint angles as the seated cycle condition. Within their transitions were multiple smaller peaks and troughs. The range of motion for standing cycling was 92.3 deg. It began at 160.7 deg, flexed to 74.1 deg, and extended again to 159.4 deg. The run condition began with an extended knee angle of 166.5 deg, maximally flexed to 78.3 deg, and ended in an extended position of 158.9 deg. The range of motion of the knee during running was 82.5 deg (Figure 2a, Table 2).
Hip
Both cycle conditions followed the same general trend of beginning in maximum hip extension, moving toward maximum hip flexion, and then moving back to maximum hip extension. Specifically, the seated cycle condition began at a maximum extension of 139.5 deg, flexed to 86.6 deg, and extended again to 126.9 deg. The hip achieved a range of motion of 43.9 deg during seated cycling. The standing cycle condition began maximally extended at 130.8 deg, flexed to 91.3 deg, and then extended again to 125.8 deg. The hip achieved a range of motion of 39.0 deg during standing cycling (Figure 2b, Table 2).
The run condition is characterized by two hip extension peaks with a small flexion trough in between. The sequence of run hip joint angles was an initial angle of 140.8 deg that extended to the first peak angle of 190.9 deg, followed by flexion to 131.5 deg, which then extended again to the second peak angle of 162.1 deg. The hip finally moved through more flexion ending at an angle of 148.6 deg. The hip achieved a range of motion of 54.9 deg during running (Figure 2b, Table 2).
Ankle
Again, both cycle conditions followed the same general trend and exhibited a wavelike pattern of peaks and troughs throughout the ankle joint angle measurements with greater plantarflexion crests in the beginning of the period followed by a dip in extension measures, or more dorsiflexion angle measurements in the second half of the period. Specifically, the standing cycle condition reached a maximum plantarflexion angle of 141.1 deg in the first half of the movement and a maximum dorsiflexion angle of 96.4 deg in the second half of the movement. The ankle obtained a range of motion of 44.7 deg. In the seated cycle condition, a maximum plantarflexion angle of 124.3 deg was observed in the first half of the movement, which was followed by a maximum dorsiflexion angle of 94.7 deg in the second half of the movement. The range of motion of the ankle during seated cycling was 29.6 deg (Figure 2c, Table 2).
Once more the run condition ankle joint measurements followed an independent trend. Rather than showing plantarflexion followed by dorsiflexion in small wave increments as did the cycle conditions, the run condition exhibited a more definite peak and valley pattern of a dorsiflexion valley followed by a plantarflexion peak. It ended in a major dorsiflexion drop that worked its way back to plantarflexion. Specifically, the ankle began to plantarflex at 117.1 deg, dorsiflexed down to 96.2 deg, plantarflexed back to 135.3 deg, and then dorsiflexed to 53.3 deg. The range of motion of the ankle during running was 82.0 deg (Figure 2c, Table 2).
Joint Angular Velocities
Knee
The run and standing cycling conditions followed the same trend of having multiple velocity peaks, whereas seated cycle exhibited a single wave pattern. The seated condition began with a -91.7 deg/s velocity, dipped to -362.8 deg/s, climbed to 334.9 deg/s, and finished at 81.9 deg/s. Standing cycle velocity began at 72.4 deg/s, dropped to -406.3 deg/s, showed two dramatic peak velocities of 539.1 deg/s and 487.3 deg/s, and finally ended with a velocity of 108.1 deg/s. In the running condition, initial velocity was -228.4 deg/s, it dropped to -402.7 deg/s, increased to its first peak velocity of 206.5 deg/s, dropped again to -323.7 deg/s, and then reached a second peak velocity of 664.4 deg/s before finally ending with a velocity of -164.8 deg/s (Figure 3a, Table 3).
Hip
The two cycle conditions exhibited similar behaviors in that they oscillated through the same velocities throughout the movement. Hip angular velocities in the run condition, however, underwent dramatic changes in the first half of the movement. In the seated cycle condition, hip velocity for the first .4 s remained in the range of approximately -42 deg/s to approximately -204 deg/s, then at .4 s the velocities increased to 152.2 deg/s and remained in the range of 68.4 deg/s to 212 deg/s for the remainder of the movement. The stand cycle condition followed a wavelike pattern with small, random waves remaining in the range of -230.7 deg/s to 245 deg/s. Finally, the run condition angular velocities experienced great changes in the first .23 s, ranging from -2306.7 deg/s, up to 2256.9 deg/s, and then back to -402.6 deg/s. For the remainder of the period, hip angular velocities remained in the range of 326.4 deg/s to -229.3 deg/s (Figure 3b, Table 3).
Ankle
In terms of ankle angular velocities, all three conditions exhibited oscillations throughout the movement. Both cycle conditions had smaller and more frequent changes in ankle angular velocities, whereas the run condition demonstrated three exaggerated changes in direction of the velocities. The seated cycle condition angular velocities fluctuated within the range of -199.2 deg/s and 176.6 deg/s. The stand cycle condition vacillated through a greater range of -319.7 deg/s and 227.4 deg/s. The run condition initially began by dipping down to -260.7 deg/s, shot up to a peak of 510.7 deg/s, dropped again to -760.7 deg/s, peaked again at 904.7 deg/s, and then smoothed out to a range of 91.3 deg/s and -104.3 deg/s (Figure 3c, Table 3).

DISCUSSION

In this study we conclude that running is kinematically more similar to standing cycling than seated cycling. Results supporting this conclusion include joint angle measurements, knee angular velocities, and condition frequencies.
Concerning joint angle measurements, the run knee maximum and minimum joint angle measures resemble standing cycling maximum and minimum knee joint angle measures more than they do seated cycling maximum and minimum knee joint angle measures (Table 2). Minimum hip joint angle measures are more similar between running and standing cycling than between running and seated cycling (Table 2). Maximum ankle joint measures are also more similar between running and standing cycling than between running and seated cycling (Table 2).
Concerning joint angular velocities, the run knee maximum and minimum negative and positive angular velocities are more similar to standing cycling maximum and minimum negative and positive velocities than to seated cycling maximum and minimum negative and positive velocities (Table 3). Finally, the run frequency is closer in value to standing cycling than to seated cycling (Table 1).
The ranges of motions found for the lower extremity joints in each of the conditions do not, however, support our conclusion (Table 2). We postulate that this is due to the fact that running generally has a free range of motion while cyclists clip into the bike so that their range of motion is limited. A decreased range of motion may also help to explain why some of the running and standing cycling joint angles are not more similar than running and seated cycling joint angles. In this study, it could not be concluded that the hip maximum joint angle and the ankle minimum joint angle (Table 2) are more similar between running and standing cycling than between running and seated cycling because the standing cycling and seated cycling joint angles in both cases were so close in value that they were essentially the same. Thus, one is not more similar to running than the other.
Additional explanations offering support for the concept that standing cycling is more comparable to running than seated cycling involve more similar exercise intensities, more similar muscular activity and force production, and finally more similar frequencies.
Heart Rate
In this investigation and in the literature, exercise intensity, measured by heart rate, is more similar between running conditions and standing cycling than between running conditions and seated cycling. In the present study, seated cycling heart rate is significantly lower than running heart rate and standing cycling heart rate, but standing cycling heart rate is not significantly different from running heart rate (Table 1).
Accordingly, in a study of seven professional cyclists comparing heart rates during the three main competition phases in stage races including flat stages, time trials, and high mountains stages, the heart rates between flat stages and high mountain stages were significantly different. These stages were chosen for comparison because they are most applicable to the present study, in that the cycling in the flat stages is fast and level, such as the seated cycle portion, whereas the high mountain stage cycling is uphill climbing that requires a large degree of out of saddle pedaling, which is similar to the stand cycle condition. Flat terrain heart rates were 124 +/-2 bpm, and high mountain heart rates were 157+/-4 bpm (Lucía, Hoyos, & Chicarro, 2001).
In another study comparing heart rates of standing cycling to seated cycling, 10 bicycle racers from a local racing club were tested, and again it was found that standing heart rate, 137 +/- 5 bpm, was significantly different from seated cycling heart rate, 127 +/- 4 bpm (Ryschon & Stray-Gundersen, 1991). Both of these studies lend support to the idea that there is a significant difference in exercise intensity between seated and standing cycling as is found in the present study.
Expanding on this conclusion, if one of these exercise intensities, standing or seated, is more similar to a run exercise intensity, then it could be extrapolated that performing a certain cycle condition will more prepare a triathlete for the run portion of a race by matching the exercise intensity of running. In a study that had 7 triathletes perform three experimental trials which involved completing a 2 hr 15 min simulated triathlon, a 2 hr 15 min marathon with the last 45 min of the run at the same speed as a triathlon race pace and, a 45 min isolated run at triathlon race pace, heart rate was recorded for each condition. Isolated run heart rates are the only data cited here, for in the present study only separate trials of running and cycling are applicable, not data illustrating the effect of one on the other. Isolated run heart rates were 147 +/- 13.3 bpm (at 2-5 min), 151 +/- 13.1 bpm (at 15-22 min), and 154 +/- 13.5 bpm (at 35-42 min), (Hausswirth, Bigard, & Guezennec, 1997). All of these numbers are significantly different from the seated cycling heart rates cited earlier, but they are not significantly different from standing cycle heart rates cited earlier.
Thus, the literature credits the data gathered in this study by demonstrating that running heart rates are significantly different from seated cycling heart rates but not from standing cycling heart rates. One strategy to more prepare a triathlete for the run portion of a race is to have the triathlete perform standing cycling before running to mimic the different exercise intensity of running. It should be noted that the numbers collected in the present study are from one subject and thus do not represent a normal distribution. They can, however, be applied to numbers found in other studies as the heart rate values in the present study and in the literature can support one another in general ideas and concepts.
Muscle Activity and Force Production
In all three investigated conditions of seated cycling, standing cycling, and running, the same muscles of the lower extremities are used: hip extensors, gluteal muscles, quadriceps, hamstrings, gastrocnemius, soleus, and the anterior tibialis (Sanner & O'Halloran, 2000; Novacheck, 1998; Adelaar, 1986). Differences do exist, however, in the way these muscles are used. First of all, some researchers observed that more muscular activity occurs in standing cycling versus seated cycling (Ryschon & Stray-Gundersen, 1991). They speculate that this increased activity is due to a need for torso stabilization and to guide the bike in a side-to-side rocking motion, which often occurs in standing cycling. By recruiting more muscles of the upper body we suggest that standing cycling is more like running because running demands the use of upper body muscles such as those used in arm swinging, balance, and stabilization.
Another muscular difference is that cycling is characterized as concentric and nonweight-bearing, whereas running is characterized as being largely eccentric and weight-bearing (Quigley & Richards, 1996). This difference elicits a discussion of the forces involved. The literature repeatedly reports that the ground reaction forces (GRFs) in running are 2-3 times the body weight (Quigley & Richards, 1996; Novacheck, 1998; Dixon, Collop, & Batt, 2000). Also, several sources report that in standing cycling, normal forces 2-3 times body weight are generated because the rider transitions from seated cycling, a non weight-bearing activity, to standing on the pedals, a weight-bearing activity (Ryschon & Stray-Gundersen, 1991; Caldwell, Li Li, McCole, & Hagberg, 1998; Millet & Vleck, 1999).
In connection with the forces involved, it could be suggested that perhaps there is also more eccentric contraction in standing cycling because force generation is greater. One of the means to increase force generation is the stretch-shorten cycle, or eccentric contractions directly preceding concentric contractions. Therefore, the greater forces generated in standing cycling could result from having to supporting body weight and from a larger eccentric component. Both of these qualities make standing cycling more similar to running, and they give support to the idea that triathletes can prepare their bodies for the run portion of a race by subjecting their bodies to the forces and muscle activity that will be expected in the run by standing at the end of the cycle portion of a race.
Frequencies
In this study, run frequency was found to be 1.37 Hz, standing cycling frequency was found to be 1.25 Hz, and seated cycling was found to be 1.52 Hz (Table 1). The major trend in this study is that standing and seated cycling frequencies differ through a wide range, and in the case of this study, run frequency is closer in value to standing cycling than seated cycling.
The literature reports running frequencies ranging from approximately 1.0 to 1.5 Hz (Millet & Vleck, 1999; Gottschall & Palmer, 2000; Creagh, Reilly, & Lees, 1998), standing cycling frequencies of approximately 1.68 Hz (Caldwell, Hagberg, McCole, & Li Li, 1999), and seated cycling frequencies of approximately 1.5 to 2.5 Hz (Millet & Vleck, 1999; Caldwell, Hagberg, McCole, & Li Li, 1999; Marsh & Martin, 1995). The range of frequencies for seated cycling extends higher than the range of frequencies for standing cycling and for running.
Running frequencies and standing cycling frequencies are both slower than seated cycling frequencies based on the literature and results obtained in this study. Therefore, one could argue that a triathlete could more prepare their body for the run portion of the race by standing while cycling at the end of the cycle portion based on the fact that standing cycling frequencies are more similar to run frequencies than seated cycling frequencies are to run frequencies. When the athlete stands up on the bike, he slows the cadence of his legs and better prepares his legs in muscle and neural activity for the rhythm of running.
In summary, based on the kinematic results of this study, including the maximum and minimum knee joint angles, minimum hip joint angles, maximum ankle joint angles, maximum and minimum negative and positive knee angular velocities, and condition frequencies, we conclude that standing cycling is kinematically more similar to running than seated cycling. In addition, literature pertaining to exercise intensity, muscle activity and force, and condition frequencies lends further support to our conclusion that standing cycling is kinematically more similar to running than seated cycling.

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