Course Syllabus and Overview

Physics 323/623: Computational Biophysics Lab

Time:  Wed 1:30-3:30

Instructors: Drs. Jacquelyn S. Fetrow and Freddie R. Salsbury, Jr.

Office: Olin 301A and Olin 301B

Email: salsbufr@wfu.edu, fetrowjs@wfu.edu

Web page: http://www.wfu.edu/~fetrowjs/Teaching.htm

Office Hours: TBA

Textbook:  Molecular Modeling, Principles and Applications (Leach), Prentice Hall; this comprehensive textbook will provide theory and background on the methods that we will be using in this course. You will not be required to know the chapters that we do not cover; we will also use the lecture textbook, Molecular Biophysics (Duane), for supplemental reading during the semester

Research-based learning: The best way to learn to use computational biophysics methods is to apply those methods in a research-based format and we will follow this learning approach in this course.  We will teach methods and theory, but you will apply the methods and theory to a problem for which we do not yet know the “right answer;” however, it is a problem in which we are interested. 

            This semester, we have chosen the protein peroxiredoxin (Prx) for analysis.  Each of you will be assigned a different Prx structure.  This structure will be the one that you use and analyze during each of the laboratory exercises.  Your grade will be partially dependent on your knowledge of your structure, your observations and analysis of the results you obtain for your structure, and your critical comparison of your data to the results produced by other students on their own Prx structures.

Background reading for Prxs: The following journal articles will provide critical background reading for the Prx protein family.  All are available on PubMed and all are required reading.

  • Hofmann, B., Hecht, H.-J., and Flohe, L. (2002) Peroxiredoxins. Biol. Chem. 383, 347-364.
  • Wood, Z. A., Schroder, E., Harris, J. R., and Poole, L. B. (2003b) Structure, mechanism and regulation of peroxiredoxins. Trends Biochem Sci 28, 32-40.
  • Wood, Z. A., Poole, L. B., and Karplus, P. A. (2003a) Peroxiredoxin evolution and the regulation of hydrogen peroxide signaling. Science 300, 650-653

Grading:

            Labs (8 exercises, 50 points each)                                        400 points (480 gr)

            Quiz                                                                                        50 points

            Class participation, observations, and creativity                    100 points

            Paper (4 drafts, 50 points each; final paper 100 points)        300 points

            Total:                                                                                       850 points

Graduate credit:  Students registered for any of the graduate course numbers and receiving graduate credit will be held to higher expectations than students receiving undergraduate credit. Graduate students will be expected to answer lab and exam questions in more detail.  Often, there will be an additional, more difficult question that graduate students must answer in addition to the other questions.  Graduate students are expected to participate in class more often and to offer more insightful observations.

Software/hardware required:  Laptop computer, loaded with SSH, Hummingbird Exceed (X-windowing software), NAMD (DEAC cluster), VMD (DEAC cluster), MEAD (DEAC cluster), MatLab (laptop); printer access; Blackboard

 

 

Tentative Schedule

Date

Topic

Reading Assignment (Read prior to coming to lab)

Quizzes and Papers

Laboratory Assignment Due Date (due by the beginning of lab on due date)

Leader

Wed Aug 25

Lab 1: Load software; conformational analysis using physical models

Leach, Sections 1.1-1.9, 9.1-9.3, 10.1-10.2; Duane, Sections 4.1-4.4, 5.1-5.4 (Leach, 4.10)*

 

 

JSF

Wed Sep 1

Lab 2: Introduction to protein structure web sites

Section 9.11, Appendix 10.2, Leach; Prx journal articles (Duane, Section 5.6)

Quiz (amino acids and nucleotides)

Lab 1 due

JSF

Wed Sep 8

Lab 3: Introduction to DEAC cluster, Linux operating system; editing and queueing

Prx journal articles for assigned structure

Paper, chapter 1, Prx seq, struct, funct

Lab 2 due

FRS

Wed Sep15

Lab 4: Dynamics in explicit solvent

Sections 4.1-4.16, 5.1-5.7, 6.4-6.9, Leach

 

Lab 3 due

JSF, FRS

Wed Sep 22

Lab 4: Dynamics in explicit solvent

Sections 4.14, 7.1-7.9, Leach

 

 

JSF, FRS

Wed Sep 29

Lab 5: Molecular visualization using VMD

Duane, Section 5.6

 

Lab 4 due

JSF

Wed Oct 6

Lab 5 (cont): Molecular visualization using VMD

Duane, Section 5.6

Paper, chapter 1  draft 2; chapter 2 (methods) draft 1

 

JSF

Wed Oct 13

Review of simulation strategy and methods

Sections 5.1-5.7, 6.4-6.9, 7.1-7.9 Leach

 

Lab 5 due

JSF/ FRS

Wed Oct 20

Lab 6: NAMD trajectory analysis (CHARMM and VMD)

Sections 6.6, 7.1-7.9, Leach

 

 

JSF

Wed Oct 27

Lab 6: NAMD trajectory analysis (CHARMM and VMD)

Sections 6.6, 7.1-7.9, Leach

 

 

JSF

Wed Nov 3

Lab 6: NAMD trajectory analysis (comparison between student results)

Sections 6.6, 7.1-7.9, Leach

Paper, chapter 1, draft 3; chapter 2 (methods and results) draft 2

 

JSF

Wed Nov 10

Lab 7: Electrostatic calculations:  MEAD

 

 

Lab 6 due

FRS

Wed Nov 17

Lab 7: Analysis of electrostatic calculations; comparison between student computational results; final discussion of research paper

Section 4.9, Leach

Paper, chapter 2 (complete, draft 3); chapter 3 (methods) draft 1; chapter 4 (interpretation of MD results in terms of function, draft 1)

 

FRS

Wed Dec 1

Lab 7: Analysis of electrostatic calculations; comparison between student computational results; final discussion of research paper

 

 

 

FRS/ JSF

Thurs Dec 2

[Bashford seminar]

 

 

Lab 7 due

 

Mon Dec 6

 

 

Final paper due (including final chapters on electrostatics and critical analysis)

 

 

*Reading material in parentheses is not required reading, but is provided for further reference and is recommended reading prior to coming to lab.