Evolution

Biology 314

Herman Eure

Charles Bonnet, a French embryologist, coined the term evolution to explain his embryological theory called Preformation or Emboitement. His idea was that there was, in each embryo, an encapsulated miniature adult, and production of the mature adult form was accomplished by a series of developmental unfoldings. Evolution (e = out + volutio = turning or folding) thus was an unfolding or turning out of this miniature adult to create the mature form. Today we know that there is no miniature adult encapsulated in a developing embryo. However, what we do know is that each zygote carries within it a genetic map that controls its development from conception/fertilization to the production of the adult forms. Populations of these organisms carry this same map but may express it differently in a given environment. Evolution is the process of genetic change in this map through time. The mechanism that allows for the creation of new and different "types" from an "old standard" is natural selection. This course is designed to study the development of evolutionary thought beginning with the Greek Philosopher, Thales, who lived some 2000 years before Charles Darwin to Charles Darwin and finally to today’s molecular geneticists. This course looks at the history of evolutionary thought including the study of those individuals whose ideas contributed to Darwin’s synthetic theory of evolution. Aspects of historical geology, including those principles and processes that were instrumental in both Charles Darwin’s and Alfred Russel Wallace's independent discovery of the process of genetic change in populations, are explored. Mechanisms of evolution, including changes in gene frequencies, selection and adaptation, speciation, variability, geographical variation and isolation, polymorphism, structural and interaction of populations, and molecular genetics are discussed. Mechanisms that preserve variability including an analysis of the Hardy-Weinberg equilibrium are explored. Finally the course looks at evolutionary lineages, phylogenies, extinction and causes of extinction. The course has one lab class meeting where students can look at fossils to augment the information that they have been discussing in class. The class is taught each fall and is open to sophomores, juniors and seniors.